| Class a: All alpha proteins [46456] (284 folds) | 
| Fold a.22: Histone-fold [47112] (1 superfamily) core: 3 helices; long middle helix is flanked at each end with shorter ones  | 
Superfamily a.22.1: Histone-fold [47113] (4 families) ![]()  | 
| Family a.22.1.1: Nucleosome core histones [47114] (6 proteins) form octamers composed of two copies of each of the four histones  | 
| Protein Histone H3 [47122] (5 species) | 
| African clawed frog (Xenopus laevis) [TaxId:8355] [47124] (38 PDB entries) | 
| Domain d1aoia_: 1aoi A: [16462] Other proteins in same PDB: d1aoib_, d1aoic_, d1aoid_, d1aoif_, d1aoig_, d1aoih_ protein/DNA complex; complexed with mn  | 
| Domain d1aoie_: 1aoi E: [16463] Other proteins in same PDB: d1aoib_, d1aoic_, d1aoid_, d1aoif_, d1aoig_, d1aoih_ protein/DNA complex; complexed with mn  | 
| Domain d1f66a_: 1f66 A: [16460] Other proteins in same PDB: d1f66b_, d1f66c_, d1f66d_, d1f66f_, d1f66g_, d1f66h_ protein/DNA complex; complexed with mn  | 
| Domain d1f66e_: 1f66 E: [16461] Other proteins in same PDB: d1f66b_, d1f66c_, d1f66d_, d1f66f_, d1f66g_, d1f66h_ protein/DNA complex; complexed with mn  | 
| Domain d1kx3a_: 1kx3 A: [77577] Other proteins in same PDB: d1kx3b_, d1kx3c_, d1kx3d_, d1kx3f_, d1kx3g_, d1kx3h_ protein/DNA complex; complexed with mn  | 
| Domain d1kx3e_: 1kx3 E: [77581] Other proteins in same PDB: d1kx3b_, d1kx3c_, d1kx3d_, d1kx3f_, d1kx3g_, d1kx3h_ protein/DNA complex; complexed with mn  | 
| Domain d1kx4a_: 1kx4 A: [77585] Other proteins in same PDB: d1kx4b_, d1kx4c_, d1kx4d_, d1kx4f_, d1kx4g_, d1kx4h_ protein/DNA complex; complexed with cl, mn  | 
| Domain d1kx4e_: 1kx4 E: [77589] Other proteins in same PDB: d1kx4b_, d1kx4c_, d1kx4d_, d1kx4f_, d1kx4g_, d1kx4h_ protein/DNA complex; complexed with cl, mn  | 
| Domain d1kx5a_: 1kx5 A: [77593] Other proteins in same PDB: d1kx5b_, d1kx5c_, d1kx5d_, d1kx5f_, d1kx5g_, d1kx5h_ protein/DNA complex; complexed with cl, mn  | 
| Domain d1kx5e_: 1kx5 E: [77597] Other proteins in same PDB: d1kx5b_, d1kx5c_, d1kx5d_, d1kx5f_, d1kx5g_, d1kx5h_ protein/DNA complex; complexed with cl, mn  | 
| Domain d1m18a_: 1m18 A: [78371] Other proteins in same PDB: d1m18b_, d1m18c_, d1m18d_, d1m18f_, d1m18g_, d1m18h_ protein/DNA complex; complexed with abu, imt, mn  | 
| Domain d1m18e_: 1m18 E: [78375] Other proteins in same PDB: d1m18b_, d1m18c_, d1m18d_, d1m18f_, d1m18g_, d1m18h_ protein/DNA complex; complexed with abu, imt, mn  | 
| Domain d1m19a_: 1m19 A: [78379] Other proteins in same PDB: d1m19b_, d1m19c_, d1m19d_, d1m19f_, d1m19g_, d1m19h_ protein/DNA complex; complexed with imt, mn  | 
| Domain d1m19e_: 1m19 E: [78383] Other proteins in same PDB: d1m19b_, d1m19c_, d1m19d_, d1m19f_, d1m19g_, d1m19h_ protein/DNA complex; complexed with imt, mn  | 
| Domain d1m1aa_: 1m1a A: [78387] Other proteins in same PDB: d1m1ab_, d1m1ac_, d1m1ad_, d1m1af_, d1m1ag_, d1m1ah_ protein/DNA complex; complexed with imt, mn  | 
| Domain d1m1ae_: 1m1a E: [78391] Other proteins in same PDB: d1m1ab_, d1m1ac_, d1m1ad_, d1m1af_, d1m1ag_, d1m1ah_ protein/DNA complex; complexed with imt, mn  | 
| Domain d1s32a_: 1s32 A: [98412] Other proteins in same PDB: d1s32b_, d1s32c_, d1s32d_, d1s32f_, d1s32g_, d1s32h_ protein/DNA complex; complexed with cl, imt, mn, ogg  | 
| Domain d1s32e_: 1s32 E: [98416] Other proteins in same PDB: d1s32b_, d1s32c_, d1s32d_, d1s32f_, d1s32g_, d1s32h_ protein/DNA complex; complexed with cl, imt, mn, ogg  | 
| Domain d1zbba1: 1zbb A:39-135 [145985] Other proteins in same PDB: d1zbbb1, d1zbbd1, d1zbbf1 automatically matched to d1p3ie_ protein/DNA complex  | 
| Domain d1zbbe1: 1zbb E:39-135 [145988] Other proteins in same PDB: d1zbbb1, d1zbbd1, d1zbbf1 automatically matched to d1p3ie_ protein/DNA complex  | 
| Domain d1zlaa_: 1zla A: [125240] Other proteins in same PDB: d1zlab1, d1zlac1, d1zlad1, d1zlaf1, d1zlag1, d1zlah1 automated match to d1kx5a_ protein/DNA complex  | 
| Domain d1zlae_: 1zla E: [125244] Other proteins in same PDB: d1zlab1, d1zlac1, d1zlad1, d1zlaf1, d1zlag1, d1zlah1 automated match to d1kx5a_ protein/DNA complex  | 
| Domain d2fj7a1: 2fj7 A:38-135 [133548] Other proteins in same PDB: d2fj7b1, d2fj7d1, d2fj7f1, d2fj7h1 automatically matched to d1p3ie_ protein/DNA complex  | 
| Domain d2fj7e1: 2fj7 E:38-135 [133551] Other proteins in same PDB: d2fj7b1, d2fj7d1, d2fj7f1, d2fj7h1 automatically matched to d1p3ie_ protein/DNA complex  | 
| Domain d2io5b_: 2io5 B: [161465] Other proteins in same PDB: d2io5a1, d2io5c1 automated match to d1kx5a_  | 
| Domain d2nzda_: 2nzd A: [138848] Other proteins in same PDB: d2nzdb1, d2nzdc_, d2nzdf1, d2nzdg_ automated match to d1kx5a_ protein/DNA complex; complexed with mn  | 
| Domain d2nzde_: 2nzd E: [138851] Other proteins in same PDB: d2nzdb1, d2nzdc_, d2nzdf1, d2nzdg_ automated match to d1kx5a_ protein/DNA complex; complexed with mn  | 
| Domain d3b6fa1: 3b6f A:33-135 [154884] Other proteins in same PDB: d3b6fb1, d3b6fc1, d3b6fd1, d3b6ff1, d3b6fg1, d3b6fh1 automatically matched to d1aoie_ protein/DNA complex; complexed with mn  | 
| Domain d3b6fe1: 3b6f E:33-135 [154888] Other proteins in same PDB: d3b6fb1, d3b6fc1, d3b6fd1, d3b6ff1, d3b6fg1, d3b6fh1 automatically matched to d1aoie_ protein/DNA complex; complexed with mn  | 
| Domain d3b6ga1: 3b6g A:33-135 [154892] Other proteins in same PDB: d3b6gb1, d3b6gc1, d3b6gd1, d3b6gf1, d3b6gg1, d3b6gh1 automatically matched to d1aoie_ protein/DNA complex; complexed with mn  | 
| Domain d3b6ge1: 3b6g E:33-135 [154896] Other proteins in same PDB: d3b6gb1, d3b6gc1, d3b6gd1, d3b6gf1, d3b6gg1, d3b6gh1 automatically matched to d1aoie_ protein/DNA complex; complexed with mn  | 
| Domain d3c1ba_: 3c1b A: [155851] Other proteins in same PDB: d3c1bb_, d3c1bc1, d3c1bd1, d3c1bf_, d3c1bg1, d3c1bh1 automated match to d1kx5a_ protein/DNA complex  | 
| Domain d3c1be_: 3c1b E: [155855] Other proteins in same PDB: d3c1bb_, d3c1bc1, d3c1bd1, d3c1bf_, d3c1bg1, d3c1bh1 automated match to d1kx5a_ protein/DNA complex  | 
| Domain d3kuya_: 3kuy A: [179728] Other proteins in same PDB: d3kuyc_, d3kuyg_ automated match to d1kx5a_ protein/DNA complex; complexed with atv, mn  | 
| Domain d3kuye_: 3kuy E: [179730] Other proteins in same PDB: d3kuyc_, d3kuyg_ automated match to d1kx5a_ protein/DNA complex; complexed with atv, mn  | 
| Domain d3lela_: 3lel A: [180229] automated match to d1kx5a_ protein/DNA complex; complexed with mn  | 
| Domain d3lele_: 3lel E: [180230] automated match to d1kx5a_ protein/DNA complex; complexed with mn  | 
| Domain d3lelk_: 3lel K: [180231] automated match to d1kx5a_ protein/DNA complex; complexed with mn  | 
| Domain d3lelo_: 3lel O: [180232] automated match to d1kx5a_ protein/DNA complex; complexed with mn  | 
| Domain d3ljaa_: 3lja A: [180318] automated match to d1kx5a_ protein/DNA complex; complexed with mn, so4  | 
| Domain d3ljae_: 3lja E: [180319] automated match to d1kx5a_ protein/DNA complex; complexed with mn, so4  | 
| Domain d3lz0a_: 3lz0 A: [180643] Other proteins in same PDB: d3lz0c_, d3lz0g_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, mn  | 
| Domain d3lz0e_: 3lz0 E: [180645] Other proteins in same PDB: d3lz0c_, d3lz0g_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, mn  | 
| Domain d3lz1a_: 3lz1 A: [180647] Other proteins in same PDB: d3lz1c_, d3lz1g_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, mn  | 
| Domain d3lz1e_: 3lz1 E: [180649] Other proteins in same PDB: d3lz1c_, d3lz1g_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, mn  | 
| Domain d3mgpa_: 3mgp A: [181193] Other proteins in same PDB: d3mgpc_, d3mgpg_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, co  | 
| Domain d3mgpe_: 3mgp E: [181195] Other proteins in same PDB: d3mgpc_, d3mgpg_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, co  | 
| Domain d3mgqa_: 3mgq A: [181197] Other proteins in same PDB: d3mgqc_, d3mgqg_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, ni  | 
| Domain d3mgqe_: 3mgq E: [181199] Other proteins in same PDB: d3mgqc_, d3mgqg_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, ni  | 
| Domain d3mgra_: 3mgr A: [181201] Other proteins in same PDB: d3mgrc_, d3mgrg_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, mn, rb  | 
| Domain d3mgre_: 3mgr E: [181203] Other proteins in same PDB: d3mgrc_, d3mgrg_ automated match to d1kx5a_ protein/DNA complex; complexed with cl, mn, rb  | 
| Domain d3mnna_: 3mnn A: [181451] Other proteins in same PDB: d3mnnc_, d3mnng_ automated match to d1kx5a_ protein/DNA complex; complexed with mg, mml, ptw, ru, so4  | 
| Domain d3mnne_: 3mnn E: [181453] Other proteins in same PDB: d3mnnc_, d3mnng_ automated match to d1kx5a_ protein/DNA complex; complexed with mg, mml, ptw, ru, so4  | 
| Domain d3mvda_: 3mvd A: [181592] automated match to d1kx5a_ protein/DNA complex  | 
| Domain d3mvde_: 3mvd E: [181593] automated match to d1kx5a_ protein/DNA complex  | 
| Baker's yeast (Saccharomyces cerevisiae) [TaxId:4932] [68986] (1 PDB entry) | 
| Domain d1id3a_: 1id3 A: [66112] Other proteins in same PDB: d1id3b_, d1id3c_, d1id3d_, d1id3f_, d1id3g_, d1id3h_ protein/DNA complex; complexed with mn  | 
| Domain d1id3e_: 1id3 E: [66116] Other proteins in same PDB: d1id3b_, d1id3c_, d1id3d_, d1id3f_, d1id3g_, d1id3h_ protein/DNA complex; complexed with mn  | 
| Chicken (Gallus gallus), erythrocytes [TaxId:9031] [47123] (5 PDB entries) Uniprot P84229  | 
| Domain d1eqzc_: 1eqz C: [16456] Other proteins in same PDB: d1eqza_, d1eqzb_, d1eqzd_, d1eqze_, d1eqzf_, d1eqzh_ protein/DNA complex; complexed with cac, cl, k, mn  | 
| Domain d1eqzg_: 1eqz G: [16457] Other proteins in same PDB: d1eqza_, d1eqzb_, d1eqzd_, d1eqze_, d1eqzf_, d1eqzh_ protein/DNA complex; complexed with cac, cl, k, mn  | 
| Domain d1hioc_: 1hio C: [16459] Other proteins in same PDB: d1hioa_, d1hiob_, d1hiod_  | 
| Domain d2hioc_: 2hio C: [16458] Other proteins in same PDB: d2hioa_, d2hiob_, d2hiod_  | 
| Fruit fly (Drosophila melanogaster) [TaxId:7227] [158386] (1 PDB entry) | 
| Domain d2pyoa_: 2pyo A: [149949] Other proteins in same PDB: d2pyob1, d2pyof1 automated match to d1kx5a_ protein/DNA complex; complexed with cl, mn  | 
| Domain d2pyoe_: 2pyo E: [149951] Other proteins in same PDB: d2pyob1, d2pyof1 automated match to d1kx5a_ protein/DNA complex; complexed with cl, mn  | 
| Mouse (Mus musculus), H3.1 [TaxId:10090] [140395] (1 PDB entry) Uniprot P68433 38-135  | 
| Domain d1u35a1: 1u35 A:438-535 [119493] Other proteins in same PDB: d1u35b1, d1u35c1, d1u35d1, d1u35f1, d1u35g1, d1u35h1 protein/DNA complex  | 
| Domain d1u35e1: 1u35 E:638-735 [119497] Other proteins in same PDB: d1u35b1, d1u35c1, d1u35d1, d1u35f1, d1u35g1, d1u35h1 automatically matched to 1U35 A:438-535 protein/DNA complex  | 
Timeline for Protein Histone H3 from a.22.1.1: Nucleosome core histones: